1. (noun) pen made from a bird's feather

• quill pen

2. (noun) any of the larger wing or tail feathers of a bird

• flight feather
• pinion
• quill feather

3. (noun) the hollow spine of a feather

• calamus
• shaft

1. (noun) pen made from a bird's feather

• pen

2. (noun) a stiff hollow protective spine on a porcupine or hedgehog

• spine

3. (noun) any of the larger wing or tail feathers of a bird

• feather
• plumage
• plume

4. (noun) the hollow spine of a feather

• rib

1. (noun) any of the larger wing or tail feathers of a bird

• primary
• primary feather
• primary quill
• tail feather

1. (noun) any of the larger wing or tail feathers of a bird

• pinion